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Joan E. Roughgarden (b. Jonathan Roughgarden in Paterson, New Jersey on 13 March 1946) is an American biologist.

Contents 1 Career 2 Selected bibliography 3 References 4 External links 5 See also

Roughgarden received a Bachelor of Science in biology and a Bachelor of Arts in Philosophy from University of Rochester in 1968 and a Ph.D. in biology from Harvard University in 1971. She is the author of 5 books and over 120 articles.

She came out as a transsexual woman at age 52 ^[1].

Roughgarden has taught at Stanford University since 1972. She founded and directed the Earth Systems Program at Stanford and was awarded for service to undergraduate education. Roughgarden's current research links ecology with economic theory.

In addition to a seminal ecology textbook written with Paul R. Ehrlich, Roughgarden published a 2004 challenge to certain tenets of sexual selection titled Evolution's Rainbow (2004): this is not an attack on the central idea of natural selection, so it is of no help to creationists; it is a correction to Darwin's explanation of the mechanism of selection. This work also contains a literature survey on unexpected sexual behavior in many species of animals: homosexual behavior; several genders and two sexes in a species (in which a gender that never participates directly in reproduction still participates indirectly, thus affecting natural selection.)

Roughgarden is also a Christian who has written on the relationship between Christianity and science ^[2].

• Roughgarden, Joan. Evolution and Christian Faith: Reflections on an Evolutionary Biologist. Hardcover ed. Washington, D.C.: Island Pr., 2006.

• Roughgarden, Joan. Evolution's Rainbow: Diversity, Gender and Sexuality in Nature and People. Paperback ed. Los Angeles: Univ. of California Pr., 2004.

• Roughgarden, Joan. Primer of Ecological Theory. 1st ed. Prentice Hall, 1997.

• Roughgarden, Jonathan. Anolis Lizards of the Caribbean: Ecology, Evolution and Plate Tectonics. Hardcover ed. Oxford Univ. Pr., 1995.

• Roughgarden, Jonathan, Robert M. May, and Simon A. Levin, eds. Perspectives in Ecological Theory. Oxford Univ. Pr., 1995.

• Ehrlich, Paul R., and Jonathan Roughgarden. Science of Ecology. Hardcover ed. Prentice Hall, 1987.

• Roughgarden, Joan. Theory of Population Genetics and Evolutionary Ecology: an Introduction. 1st ed. Prentice Hall, 1998.

1. ^ Joan Roughgarden profile: A plea for diversity. Retrieved on September 19, 2006.
2. ^ First Congregational Church of Berkeley: Event Details. Retrieved on September 19, 2006.

• Joan Roughgarden profile - Stanford Univ.

• Joan Roughgarden 50-min interv. - Web Radio (Gender Talk)

• Seed Magazine article on sexual selection theory

• Stanford University
• Against Nature?